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Monday, March 2, 2009

Fruits of the Shrubs

By Robert Adrian

Buds are young undeveloped shoots containing leaves and sometimes flowers. They are an important means of identification in winter when trees and shrubs are leafless. The terminal bud is located at the tip of a twig or branch and is usually solitary (in most species of shrubs), though there may also he two, e.g. in the lilac and bladdernut. Lateral buds are formed in the axils of the leaves and are alternately arranged, e.g. in the rose, currant, willow, or opposite, i.e. paired on either side of the twig, as in the common elder, cornelian cherry,'rivet, etc. In some species these buds are not always exactly pposite each other and are called subopposite, e.g. the spindle ree, privet and buckthorn. Flower buds vary greatly in shape, especially in the case of early flowering species such as cornelian cherry, Japanese quince, blackthorn and sea buckthorn.

The seed consists of a membranous or hard covering and inner nucleus. It is the spot through which the seed absorbs the greatest amount of water during germination and also through which the sprouts generally grow.

Only in rare instances are buds without scales and covered only with a thick pubescence, e.g. those of the wayfaring tree and alder buckthorn. These are called naked buds. Buds with scales covering only the bottom part and with the leaf tips showing are called semi-naked, e. g. those of the common elder and cotoneaster.

Other shrubs extend far to the north but do not occur in southern Europe with its warm climate. Such typical northern species are the European black currant and willow spiraea.

Still other species, e.g. the rhododendrons, green alder and rock currant, grow in the high mountain climate of central and western Europe but never occur in the north polar regions. Some shrubs grow mainly in the part of western Europe with a constant mild oceanic climate but are absent in the continental areas with severe winter frosts. These include the English holly, hawthorn and common broom.

Hypogeal germination is more common in trees with large seeds; in the case of shrubs that are found only in Corylus avellana, Rhamnus frangula and Daphne mezereum. All other shrubs are characterized by epigeal germination.

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